OK, if Behr made the tanks, who else do they make tanks for? Triumph? BMW? My guess would be yes.
You'd need to find a Triumph or BMW model that has left and right caps to verify. Then again, that would be a drastically reduced number of aftermarket caps to choose from.
If somebody installed the pop-up caps, that still suggests they made the Harley threads work. It could be done. Heck, it could be done with a little ball-burr and a Dremel if you had a steady enough hand to trim the threads down in the plastic cap.
No worries. More to come...